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Stand Structure of Northern Taiga Pine Forests

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I.N. Kutyavin, A.V. Manov, A.F. Osipov, M.A. Kuznetsov

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In Northern taiga conditions pine forests occupy about 1.8 mln ha of the territory of the Komi Republic and are mainly represented by sphagnum, green-moss and lichen forest types. The studies were carried out at the Zelenoborsk Forest Station of the Institute of Biology of the Komi Scientific Centre of the Ural Branch of the Russian Academy of Sciences in green-moss and sphagnum pine forests developing at the site of fires and clearcuts. Tree layer has a mix composition with the predominance of pine trees and admixture of spruce, birch, larch and aspen. Pine stands form three types of age structure: conventionally evenaged; conventionally uneven-aged, represented by one generation; and stepped uneven-aged, consisting of two or three generations of pine trees. Trees left behind during clearcut and felling residuals at the cutting area by the burn method contribute to the formation of stands of stepped uneven-aged age structure. The variation coefficients of tree age are 5.8–10.8 % in pine forests with one generation and 39.7–45.6 % in forests with two or three generations. The number of generations and the type of age structure had no effect on the diameter and height of trees in the stand, which varied respectively within 25.2–49.5 % and 15.7–27.8 %. In pine forests developing after fires, the peak of pine settlement occur in the second or third decade, while in cutting areas, the maximum renewal occurs in the first or second decade after clearcuts. The amplitude of tree age fluctuations varies from 16 to 33 years in conventionally even-aged and conventionally uneven-aged forests developed after clearcuts and from 30 to 45 years in post-fire stands. In stepped uneven-aged stands tree age fluctuations changes from 120 to 167 years, and from 22 to 66 years in generations. The curves showing the distribution of trees by diameter in stands of pine trees had a left-handed shift, the asymmetry is represented by positive values and varies from 0.05 to 1.03. Trees of associate species are inferior in development, both in diameter and in height to pine trees of similar age. A close correlation between the diameter and height of trees in the stands was revealed. The relationship of age with diameter and height of trees varies from weak to significant at almost all of the sites studied. A close correlation between age and diameter was found in stepped uneven-aged pine forests (R = 0.79–0.96). The correlation between diameter and age increase with an increase in tree age fluctuations amplitude.


Ivan N. Kutyavin, Candidate of Agriculture , Research Scientist; ResearcherID: P-9829-2015,
Aleksey V. Manov, Candidate of Agriculture, Research Scientist; ResearcherID: P-9089-2015,
Andrey F. Osipov, Candidate of Biology, Research Scientist; ResearcherID: P-9583-2015,
Mikhail A. Kuznetsov, Candidate of Biology, Research Scientist; ResearcherID: P-9870-2015,


Institute of Biology of Komi Science Centre of the Ural Branch of the Russian Academy of Sciences, GSP-2, ul. Kommunisticheskaya, 28, Syktyvkar, 167982, Russian Federation;


Northern taiga, pine forests, composition, structure, clearcuts, forest fires

For citation

Kutyavin I.N., Manov A.V., Osipov A.F., Kuznetsov M.A. Stand Structure of Northern Taiga Pine Forests. Lesnoy Zhurnal [Russian Forestry Journal], 2021, no. 2, pp. 86–105. DOI: 10.37482/0536-1036-2021-2-86-105


  1. Bakhtin A.A., Sokolov N.N. Types of Age Structure of Waterlogged Pine Forests in the Arkhangelsk Region. Lesnoy Zurnal [Russian Forestry Journal], 2015, no. 4, pp. 76–86. DOI: 10.17238/issn0536-1036.2015.4.76:
  2. Berezovskaya F.S., Karev G.P. Modeling of Forest Dynamics. Sibirskij Lesnoj Zurnal [Siberian Journal of Forest Science], 2015, no. 3, pp. 7–19 DOI: 10.15372/SJFS20150302
  3. Bobkova K.S., Manov A.V., Osipov V.M., Osipov A.F., Kuznetsov M.A., Torlopova N.V., Fedorkov A.L., Komarov A.S., Shanin V.N., Goncharova N.N., Zagirova S.V., Mikhaylov O.A., Miglovets M.N. Carbon in Forest and Bog Ecosystems of Specially Protected Natural Areas of the Komi Republic. Syktyvkar, 2014. 201 p.
  4. Gusev I.I. Ecosystem Modeling. Arkhangelsk, ASTU Publ., 2002. 112 p.
  5. Demakov Y.P. The Structure and Patterns of Forest Development of the Mari El Republic. Yoshkar-Ola, Volga Tech Publ., 2018. 432 p.
  6. Ermolenko P.M. Pine Forests of the Eastern Sayan. Krasnoyarsk, ILiD Publ., 1987. 148 p.
  7. Zyabchenko S.S. Pine Forests of the European North. Leningrad, Nauka Publ., 1984. 224 p.
  8. Komin G.E., Semechkin I.V. Age Structure of Forest Stands and Principles of Its Typing. Lesovedenie [Russian Journal of Forest Science], 1970, no. 2, pp. 24–33.
  9. Kuz’michev V.V. Patterns of the Dynamics of Forest Stands: Principles and Models. Novosibirsk, Nauka Publ., 2013. 208 p.
  10. Kutyavin I.N. Pine Forests of the Northern Cis-Urals: Structure, Growth, Productivity. Syktyvkar, IB FRC Komi SC UB RAS Publ., 2018. 176 p. DOI: 10.31140/book-2018-02
  11. Levin V.I. Pine Forests of the European North (Structure, Growth, and Valuation of Stands). Moscow, Lesnaya promyshlennost’ Publ., 1966. 152 p.
  12. Forest Valuation Handbook for the North-East Part of the Russian Federation (Official Materials for the Nenets Autonomous Okrug, Arkhangelsk and Vologda Regions and Komi Republic). Content by G.S. Voynov, N.P. Chuprov, S.V. Yaroslavtsev. Arkhangelsk, Pravda Severa Publ., 2012. 672 p.
  13. Listov A.A. Lichen Pine Forests. Moscow, Agropromizdat Publ., 1986. 181 p.
  14. L’vov P.N., Ipatov L.F., Plokhov A.A. Forest-Forming Processes and Their Regulation in the European North. Moscow, Lesnya promyshlennost’ Publ., 1980. 112 p.
  15. Melekhov I.S. Felling and Reforestation in the North. Arkhangelsk, Arkhangel’skoye knizhnoye izdatel’stvo, 1960. 201 p.
  16. All-Union Standards for Forest Inventory. Ed. by V.V. Zagreyev, V.I. Sukhikh, A.Z. Shvidenko, N.N. Gusev, A.G. Moshkalev. Moscow, Kolos Publ., 1992. 495 p.
  17. Komi Republic: Encyclopedia. Vol. 2. Syktyvkar, Komi knizhnoye izdatel’stvo, 1999. 573 p.
  18. Rysin L.P., Vakurov A.D., Pavlov V.F. The Value of Permanent Trial Plots in Forestry Research. Lesovedenie [Russian Journal of Forest Science], 1981, no. 1, pp. 60–66.
  19. Semenov B.A., Tsvetkov V.F., Chibisov G.A., Elizarov F.P. Trans-Tundra Forests of the European Part of Russia (Nature and Management). Arkhangelsk, Press A Publ., 1998. 334 p.
  20. Storozhenko V.G. Stable Forests Communities. The Theory and Experiment. Tula, Grif i K Publ., 2007. 192 p.
  21. Tsvetkov V.F. Pine Forests of the Kola Forest Region and Management in Them. Arkhangelsk, ASTU Publ., 2002. 380 p.
  22. Shvidenko A.Z., Schepaschenko D.G., Kraxner F., Onuchin A.A. Transition to Sustainable Forest Management in Russia: Theoretical and Methodological Backgrounds. Sibirskij Lesnoj Zurnal [Siberian Journal of Forest Science], 2017, no. 6, pp. 3–25. DOI: 10.15372/SJFS20170601
  23. Shiyatov S.G., Vaganov E.A., Kirdyanov A.V., Kruglov V.B., Mazepa V.S., Naurzbayev M.M., Khantemirov R.M. Methods of Dendrochronology. Part I. Fundamentals of Dendrochronology. Collection and Obtaining of Tree-Ring Information. Krasnoyarsk, KraSU Publ., 2000. 80 p.
  24. Del Río M., Pretzsch H., Alberdi I., Bielak B., Bravo F., Brunner A. et al. Characterization of the Structure, Dynamics, and Productivity of Mixed-Species Stands: Review and Perspectives. European Journal of Forest Research, 2016, vol. 135, iss. 1, pp. 23–49. DOI: 10.1007/s10342-015-0927-6
  25. Engelmark O., Kullman L., Bergeron Y. Fire and Age Structure of Scots Pine and Norway Spruce in Northern Sweden during the Past 700 Years. New Phytologist, 1994, vol. 126, iss. 1, pp. 163–168. DOI: 10.1111/j.1469-8137.1994.tb07542.x
  26. Garet J., Raulier F., Pothier D., Cumming S.G. Forest Age Class Structures as Indicators of Sustainability in Boreal Forest: Are We Measuring Them Correctly? Ecological Indicators, 2012, vol. 23, pp. 202–210. DOI: 10.1016/j.ecolind.2012.03.032
  27. Kuuluvainen T., Mäki J., Karjalainen L., Lehtonen H. Tree Age Distributions in Old-Growth Forest Sites in Vienansalo Wilderness, Eastern Fennoscandia. Silva Fennica, 2002, vol. 36, no. 1, pp. 169–184. DOI: 10.14214/sf.556
  28. Lilja S., Kuuluvainen T. Structure of Old Pinus sylvestris Dominated Forest Stands along a Geographic and Human Impact Gradient in Mid-Boreal Fennoscandia. Silva Fennica, 2005, vol. 39, no. 3, pp. 407–428. DOI: 10.14214/sf.377
  29. Shanin V., Komarov A., Mäkipää R. Tree Species Composition Affects Productivity and Carbon Dynamics of Different Site Types in Boreal Forests. European Journal of Forest Research, 2014, vol. 133, iss. 2, pp. 273–286. DOI: 10.1007/s10342-013-0759-1
  30. Steijlen I., Zackrisson O. Long-Term Regeneration Dynamics and Successional Trends in a Northern Swedish Coniferous Forest Stand. Canadian Journal of Botany, 1987, vol. 65, no. 5, pp. 839–848. DOI: 10.1139/b87-114
  31. Stinson G., Kurz W.A., Smyth C.E., Neilson E.T., Dymond C.C., Metsaranta J.M., Boisvenue C., Rampley G.J., Li Q., White T.M., Blain D. An Inventory-Based Analysis of Canada’s Managed Forest Carbon Dynamics, 1990 to 2008. Global Change Biology, 2011, vol. 17, iss. 6, pp. 2227–2244. DOI: 10.1111/j.1365-2486.2010.02369.x
  32. Wallenius T. Forest Age Distribution and Traces of Past Fires in a Natural Boreal Landscape Dominated by Picea abies. Silva Fennica, 2002, vol. 36, no. 1, pp. 201–211. DOI: 10.14214/sf.558

Stand Structure of Northern Taiga Pine Forests


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